Content of Natural Science Passages - PSAT Critical Reading

"The Cell Cycle" by Joseph Ritchie (2014)

The process by which cells divide and multiply is known as the cell cycle. This cycle consists of two main phases: interphase and mitosis. Each phase consists of a series of clearly defined and observable steps. At the conclusion of the cycle, each parent cell produces two genetically identical daughter cells that may also replicate by proceeding through the cell cycle.

Roughly ninety percent of the cell cycle is spent in interphase. Interphase is comprised of three main steps: the first gap phase, the synthesis phase (also called "S phase"), and the second gap phase. The initial gap phase is a period of cellular preparation in which the cell increases in size and readies itself for DNA synthesis. In the synthesis phase, or S phase, DNA replication occurs, so that when the cell divides, each daughter cell will have the DNA necessary to function properly. In the second gap phase, the cell grows in size and prepares for cellular division in the mitotic phase. At the end of each gap phase, the cell has to pass a regulatory checkpoint to ensure that nothing is going wrong. If anything has gone wrong, the checkpoints stop the cell from proceeding through the cell cycle any further.

The next part of the cell cycle is mitosis. Mitosis is a form of cell division and is broken down into five distinct phases. During prophase, the genetic material contained in the cell’s chromatin condenses into distinct chromosomes. Prometaphase is marked by the breakdown of the cell’s nuclear envelope and the formation of centrosomes at the poles of the cell. During metaphase, the cell’s chromosomes are moved to the center of the cell. A checkpoint ensures that the chromosomes are properly aligned on the center and halts the cell cycle if any errors have occurred. In anaphase, chromosomes break apart at their center, or centromere, and sister chromatids move to opposite ends of the cell. Lastly, telophase and cytokinesis occur as nuclear membranes form to physically divide the cell into two new daughter cells. Chromosomes also unwind into loose chromatin during this part of mitosis. Cytokinesis is defined as the division of the each cell’s cytoplasm and organelles. At the conclusion of the cell cycle, two genetically identical daughter cells have formed.

The cell cycle operates by a series of checkpoints and external cues. This system of checks enables the cell to enter a state of dormancy known as the gap zero phase when conditions or other factors inhibit the cell cycle. Conversely, unregulated and uncontrolled cellular division can occur under certain circumstances. A cell in a state of uncontrolled division is known to be cancerous. Lastly, cells have the ability to mediate their own death by way of apoptosis if certain genetic or physical abnormalities exist. The cell cycle is a complex process that enables cells to replicate and proliferate under a stringent set of checks and balances that produce healthy and viable daughter cells that are each able to perform the process in the future.

What is the purpose of interphase?

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

Which of the following could classify the type of people described in the underlined sentence?

"Darwinism's Effect on Science" by Matthew Minerd (2014)

For much of the history of human thought, the sciences have studied subjects that seemed to be eternal and unchanging. Even the basic laws of the Nile’s flooding were investigated in the hopes of finding never-altering laws. Similarly, the scientific investigations of the ancient Near East and Greece into the regular laws of the stars ultimately looked for constant patterns. This overall pattern of scientific reasoning has left deep marks on the minds of almost all thinkers and found its apotheosis in modern physics. From the time of the early renaissance to the nineteenth century, physics represented the ultimate expression of scientific investigation for almost all thinkers. Its static laws appeared to be the unchanging principles of all motion and life on earth. By the nineteenth century, it had appeared that only a few details had to be “cleared up” before all science was basically known.

In many ways, this situation changed dramatically with the arrival of Darwinism. It would change even more dramatically in early twentieth-century physics as well. Darwin’s theories of evolution challenged many aspects of the “static” worldview. Even those who did not believe that a divine being created an unchanging world were shaken by the new vistas opened up to science by his studies. It had been a long-accepted inheritance of Western culture to believe that the species of living organisms were unchanging in nature. Though there might be many different kinds of creatures, the kinds themselves were not believed to change. The thesis of a universal morphing of types shattered this cosmology, replacing the old world-view with a totally new one. Among the things that had to change in light of Darwin’s work was the very view of science held by most people.

Who was most affected by the changes caused by Darwinism?

"Comparing Technologies: A Difficult Endeavor" by Matthew Minerd (2014)

Comparisons of technology are often difficult to make, not only because of the rapid pace of improvements but also because of the many new applications that are available as time progresses. If we were to consider the contemporary graphing calculator and the calculation capacities of computing machines from fifty years ago, there would be astounding improvements between these two devices. However, the improvements are not reduced merely to speed improvements. A graphing calculator also has numerous output capacities that far exceed those available much older computers, none of which had the ability to represent their output in any manner even closely resembling that of contemporary devices. Merely consider the display capacities of such a device. These enable users to input many new kinds of information, enabling design engineers to design new hardware functions to match the new means of collecting user input.

The situation is even more obvious when one considers the numerous functions performed by a modern “smartphone.” These devices are equipped with a panoply of features. With all of these new functions come many new types of computational capabilities as well. In order to process images quickly, specialized hardware must be designed and software written for it in order to ensure that there are few issues with the phone’s operation. Indeed, the whole “real time” nature of telecommunications has exerted numerous pressures on the designers of computing devices. Layers of complexity, at all levels of production and development, are required to ensure that the phone can function in a synchronous manner. Gone are the days of asynchronous processing, when the computer user entered data into a mainframe, only to wait for a period of time before the processing results were provided. Today, even the smallest of digital devices must provide seamless service for users. The effects of this requirement are almost beyond number.

What is the effect of the features found on modern “smartphones”?

Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)

As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.

As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.

Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.

Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.

Based on the passage, a purpose for the damp sand was to                     .

Adapted from Volume Four of The Natural History of Animals: The Animal Life of the World in Its Various Aspects and Relations by James Richard Ainsworth Davis (1903)

The examples of protective resemblance so far quoted are mostly permanent adaptations to one particular sort of surrounding. There are, however, numerous animals which possess the power of adjusting their color more or less rapidly so as to harmonize with a changing environment.

Some of the best known of these cases are found among those mammals and birds that inhabit countries more or less covered with snow during a part of the year. A good instance is afforded by the Irish or variable hare, which is chiefly found in Ireland and Scotland. In summer, this looks very much like an ordinary hare, though rather grayer in tint and smaller in size, but in winter it becomes white with the exception of the black tips to the ears. Investigations that have been made on the closely allied American hare seem to show that the phenomenon is due to the growth of new hairs of white hue.

The common stoat is subject to similar color change in the northern parts of its range. In summer it is of a bright reddish brown color with the exception of the under parts, which are yellowish white, and the end of the tail, which is black. But in winter, the entire coat, save only the tip of the tail, becomes white, and in that condition the animal is known as an ermine. A similar example is afforded by the weasel. The seasonal change in the vegetarian Irish hare is purely of protective character, but in such an actively carnivorous creature as a stoat or weasel, it is aggressive as well, rendering the animal inconspicuous to its prey.

Based on the passage, which of the following can we infer would be the best reason for animals living in variable arctic environments to change their fur color?

Adapted from “Feathers of Sea Birds and Wild Fowl for Bedding” from The Utility of Birds by Edward Forbush (ed. 1922)

In the colder countries of the world, the feathers and down of waterfowl have been in great demand for centuries as filling for beds and pillows. Such feathers are perfect non-conductors of heat, and beds, pillows, or coverlets filled with them represent the acme of comfort and durability. The early settlers of New England saved for such purposes the feathers and down from the thousands of wild-fowl which they killed, but as the population increased in numbers, the quantity thus furnished was insufficient, and the people sought a larger supply in the vast colonies of ducks and geese along the Labrador coast.

The manner in which the feathers and down were obtained, unlike the method practiced in Iceland, did not tend to conserve and protect the source of supply. In Iceland, the people have continued to receive for many years a considerable income by collecting eider down, but there they do not “kill the goose that lays the golden eggs.” Ducks line their nests with down plucked from their own breasts and that of the eider is particularly valuable for bedding. In Iceland, these birds are so carefully protected that they have become as tame and unsuspicious as domestic fowls In North America. Where they are constantly hunted they often conceal their nests in the midst of weeds or bushes, but in Iceland, they make their nests and deposit their eggs in holes dug for them in the sod. A supply of the ducks is maintained so that the people derive from them an annual income.

In North America, quite a different policy was pursued. The demand for feathers became so great in the New England colonies about the middle of the eighteenth century that vessels were fitted out there for the coast of Labrador for the express purpose of securing the feathers and down of wild fowl. Eider down having become valuable and these ducks being in the habit of congregating by thousands on barren islands of the Labrador coast, the birds became the victims of the ships’ crews. As the ducks molt all their primary feathers at once in July or August and are then quite incapable of flight and the young birds are unable to fly until well grown, the hunters were able to surround the helpless birds, drive them together, and kill them with clubs. Otis says that millions of wildfowl were thus destroyed and that in a few years their haunts were so broken up by this wholesale slaughter and their numbers were so diminished that feather voyages became unprofitable and were given up.

This practice, followed by the almost continual egging, clubbing, shooting, etc. by Labrador fishermen, may have been a chief factor in the extinction of the Labrador duck, that species of supposed restricted breeding range. No doubt had the eider duck been restricted in its breeding range to the islands of Labrador, it also would have been exterminated long ago.

Which of the following did NOT contribute to the success and profitability of the Labrador feather voyages?

Adapted from An Introduction to Astronomy by Forest Ray Moulton (1916 ed.)

It is doubtful if any important scientific idea ever sprang suddenly into the mind of a single man. The great intellectual movements in the world have had long periods of preparation, and often many men were groping for the same truth, without exactly seizing it, before it was fully comprehended.

The foundation on which all science rests is the principle that the universe is orderly, and that all phenomena succeed one another in harmony with invariable laws. Consequently, science was impossible until the truth of this principle was perceived, at least as applied to a limited part of nature.

The phenomena of ordinary observation, as, for example, the weather, depend on such a multitude of factors that it was not easy for men in their primitive state to discover that they occur in harmony with fixed laws. This was the age of superstition, when nature was supposed to be controlled by a great number of capricious gods whose favor could be won by childish ceremonies. Enormous experience was required to dispel such errors and to convince men that the universe is one vast organization whose changes take place in conformity with laws which they can in no way alter.

The actual dawn of science was in prehistoric times, probably in the civilizations that flourished in the valleys of the Nile and the Euphrates. In the very earliest records of these people that have come down to modern times it is found that they were acquainted with many astronomical phenomena and had coherent ideas with respect to the motions of the sun, moon, planets, and stars. It is perfectly clear from their writings that it was from their observations of the heavenly bodies that they first obtained the idea that the universe is not a chaos. Day and night were seen to succeed each other regularly, the moon was found to pass through its phases systematically, the seasons followed one another in order, and in fact the more conspicuous celestial phenomena were observed to occur in an orderly sequence. It is to the glory of astronomy that it first led men to the conclusion that law reigns in the universe.

What does this passage imply to be the effect of not understanding the causes of events?

"Darwinism's Effect on Science" by Matthew Minerd (2014)

For much of the history of human thought, the sciences have studied subjects that seemed to be eternal and unchanging. Even the basic laws of the Nile’s flooding were investigated in the hopes of finding never-altering laws. Similarly, the scientific investigations of the ancient Near East and Greece into the regular laws of the stars ultimately looked for constant patterns. This overall pattern of scientific reasoning has left deep marks on the minds of almost all thinkers and found its apotheosis in modern physics. From the time of the early renaissance to the nineteenth century, physics represented the ultimate expression of scientific investigation for almost all thinkers. Its static laws appeared to be the unchanging principles of all motion and life on earth. By the nineteenth century, it had appeared that only a few details had to be “cleared up” before all science was basically known.

In many ways, this situation changed dramatically with the arrival of Darwinism. It would change even more dramatically in early twentieth-century physics as well. Darwin’s theories of evolution challenged many aspects of the “static” worldview. Even those who did not believe that a divine being created an unchanging world were shaken by the new vistas opened up to science by his studies. It had been a long-accepted inheritance of Western culture to believe that the species of living organisms were unchanging in nature. Though there might be many different kinds of creatures, the kinds themselves were not believed to change. The thesis of a universal morphing of types shattered this cosmology, replacing the old world-view with a totally new one. Among the things that had to change in light of Darwin’s work was the very view of science held by most people.

Consider the underlined sentence. What was the new “cosmology” that arose after Darwin’s day?

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

What can we say of the effects of Copernicus’ discoveries on the reigning medieval way of looking at the world?

"Interpreting the Copernican Revolution" by Matthew Minerd (2014)

The expressions of one discipline can often alter the way that other subjects understand themselves. Among such cases are numbered the investigations of Nicolaus Copernicus. Copernicus is best known for his views concerning heliocentrism, a view which eventually obliterated many aspects of the ancient/medieval worldview, at least from the standpoint of physical science. It had always been the natural view of mankind that the earth stood at the center of the universe, a fixed point in reference to the rest of the visible bodies. The sun, stars, and planets all rotated around the earth.

With time, this viewpoint became one of the major reference points for modern life. It provided a provocative image that was used—and often abused—by many people for various purposes. For those who wished to weaken the control of religion on mankind, it was said that the heliocentric outlook proved man’s insignificance. In contrast with earlier geocentrism, heliocentrism was said to show that man is not the center of the universe. He is merely one small being in the midst of a large cosmos. However, others wished to use the “Copernican Revolution” in a very different manner. These thinkers wanted to show that there was another “recentering” that had to happen. Once upon a time, we talked about the world. Now, however, it was necessary to talk of man as the central reference point. Just as the solar system was “centered” on the sun, so too should the sciences be centered on the human person.

However, both of these approaches are fraught with problems. Those who wished to undermine the religious mindset rather misunderstood the former outlook on the solar system. The earlier geocentric mindset did not believe that the earth was the most important body in the heavens. Instead, many ancient and medieval thinkers believed that the highest “sphere” above the earth was the most important being in the physical universe. Likewise, the so-called “Copernican Revolution” in physics was different from the one applied to the human person. Copernicus’ revolution showed that the human point of view was not the center, whereas the later forms of “Copernican revolution” wished to show just the opposite.

Of course, there are many complexities in the history of such important changes in scientific outlook. Nevertheless, it is fascinating to see the wide-reaching effects of such discoveries, even when they have numerous, ambiguous effects.

What does the passage say was the overall effect of the scientific revolution implied in Copernicus' discoveries?

"The Place of Lesion Studies in Neuroscience" by Samantha Winter (2013)

It’s easy to forget that the study of neuroscience originated from non-normalized, non-statistically appraised methods like lesion studies. It’s equally easy, with the advent of sophisticated technology, to render such a method obsolete. A small group of neuroscientists today make a case for the reinstitution of lesion studies—the study of abnormal brains with damaged regions in order to better understand the brain—into the twenty-first-century cognitive neuroscience realm. Their suggestion is bold, but their argument is justified.

Cognitive neuroscientists advocate for the use of convergent methods. Many of them argue that with the limitations of our existing techniques, convergent evidence is imperative for sound research. If this is the case, why ignore a method that has potential for implying causality in a domain dominated by correlational research? Rather than advocating for a single method, neuroscientists should take their own advice and use convergent techniques. Sound research should combine a variety of techniques to examine both causal relationships and overcome the individual shortcomings of each method through the use of many.

Lesion studies are also significantly more beneficial now than they were in earlier times. Neuroimaging methods have enhanced our understanding of what contributes to the brain problems most often encountered, and more refined experiments have been developed to confirm the findings from the more unreliable lesion studies. This transformation allows lesion studies to be included alongside the other systems as a mechanism for understanding the human brain.

The primary goal of this passage is                     .

Adapted from "Recent Views as to Direct Action of Light on the Colors of Flowers and Fruits" in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The theory that the brilliant colors of flowers and fruits is due to the direct action of light has been supported by a recent writer by examples taken from the arctic instead of from the tropical flora. In the arctic regions, vegetation is excessively rapid during the short summer, and this is held to be due to the continuous action of light throughout the long summer days. “The further we advance towards the north, the more the leaves of plants increase in size as if to absorb a greater proportion of the solar rays. M. Grisebach says that during a journey in Norway he observed that the majority of deciduous trees had already, at the 60th degree of latitude, larger leaves than in Germany, while M. Ch. Martins has made a similar observation as regards the leguminous plants cultivated in Lapland.” The same writer goes on to say that all the seeds of cultivated plants acquire a deeper color the further north they are grown, white haricots becoming brown or black, and white wheat becoming brown, while the green color of all vegetation becomes more intense. The flowers also are similarly changed: those which are white or yellow in central Europe becoming red or orange in Norway. This is what occurs in the Alpine flora, and the cause is said to be the same in both—the greater intensity of the sunlight. In the one the light is more persistent, in the other more intense because it traverses a less thickness of atmosphere.

Admitting the facts as above stated to be in themselves correct, they do not by any means establish the theory founded on them; and it is curious that Grisebach, who has been quoted by this writer for the fact of the increased size of the foliage, gives a totally different explanation of the more vivid colors of Arctic flowers. He says, “We see flowers become larger and more richly colored in proportion as, by the increasing length of winter, insects become rarer, and their cooperation in the act of fecundation is exposed to more uncertain chances.” (Vegetation du Globe, col. i. p. 61—French translation.) This is the theory here adopted to explain the colors of Alpine plants, and we believe there are many facts that will show it to be the preferable one. The statement that the white and yellow flowers of temperate Europe become red or golden in the Arctic regions must we think be incorrect. By roughly tabulating the colors of the plants given by Sir Joseph Hooker as permanently Arctic, we find among fifty species with more or less conspicuous flowers, twenty-five white, twelve yellow, eight purple or blue, three lilac, and two red or pink; showing a very similar proportion of white and yellow flowers to what obtains further south.

The purpose of this passage is                       .

Adapted from "Recent Views as to Direct Action of Light on the Colors of Flowers and Fruits" in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The theory that the brilliant colors of flowers and fruits is due to the direct action of light has been supported by a recent writer by examples taken from the arctic instead of from the tropical flora. In the arctic regions, vegetation is excessively rapid during the short summer, and this is held to be due to the continuous action of light throughout the long summer days. "The further we advance towards the north, the more the leaves of plants increase in size as if to absorb a greater proportion of the solar rays. M. Grisebach says that during a journey in Norway he observed that the majority of deciduous trees had already, at the 60th degree of latitude, larger leaves than in Germany, while M. Ch. Martins has made a similar observation as regards the leguminous plants cultivated in Lapland.” The same writer goes on to say that all the seeds of cultivated plants acquire a deeper color the further north they are grown, white haricots becoming brown or black, and white wheat becoming brown, while the green color of all vegetation becomes more intense. The flowers also are similarly changed: those which are white or yellow in central Europe becoming red or orange in Norway. This is what occurs in the Alpine flora, and the cause is said to be the same in both—the greater intensity of the sunlight. In the one the light is more persistent, in the other more intense because it traverses a less thickness of atmosphere.

Admitting the facts as above stated to be in themselves correct, they do not by any means establish the theory founded on them; and it is curious that Grisebach, who has been quoted by this writer for the fact of the increased size of the foliage, gives a totally different explanation of the more vivid colors of Arctic flowers. He says, “We see flowers become larger and more richly colored in proportion as, by the increasing length of winter, insects become rarer, and their cooperation in the act of fecundation is exposed to more uncertain chances.” (Vegetation du Globe, col. i. p. 61—French translation.) This is the theory here adopted to explain the colors of Alpine plants, and we believe there are many facts that will show it to be the preferable one. The statement that the white and yellow flowers of temperate Europe become red or golden in the Arctic regions must we think be incorrect. By roughly tabulating the colors of the plants given by Sir Joseph Hooker as permanently Arctic, we find among fifty species with more or less conspicuous flowers, twenty-five white, twelve yellow, eight purple or blue, three lilac, and two red or pink; showing a very similar proportion of white and yellow flowers to what obtains further south.

The author’s critique of the theory presented in the first paragraph is that                     .

Adapted from “Humming-Birds: As Illustrating the Luxuriance of Tropical Nature” in Tropical Nature, and Other Essays by Alfred Russel Wallace (1878)

The food of hummingbirds has been a matter of much controversy. All the early writers down to Buffon believed that they lived solely on the nectar of flowers, but since that time, every close observer of their habits maintains that they feed largely, and in some cases wholly, on insects. Azara observed them on the La Plata in winter taking insects out of the webs of spiders at a time and place where there were no flowers. Bullock, in Mexico, declares that he saw them catch small butterflies, and that he found many kinds of insects in their stomachs. Waterton made a similar statement. Hundreds and perhaps thousands of specimens have since been dissected by collecting naturalists, and in almost every instance their stomachs have been found full of insects, sometimes, but not generally, mixed with a proportion of honey. Many of them in fact may be seen catching gnats and other small insects just like fly-catchers, sitting on a dead twig over water, darting off for a time in the air, and then returning to the twig. Others come out just at dusk, and remain on the wing, now stationary, now darting about with the greatest rapidity, imitating in a limited space the evolutions of the goatsuckers, and evidently for the same end and purpose. Mr. Gosse also remarks, ” All the hummingbirds have more or less the habit, when in flight, of pausing in the air and throwing the body and tail into rapid and odd contortions. This is most observable in the Polytmus, from the effect that such motions have on the long feathers of the tail. That the object of these quick turns is the capture of insects, I am sure, having watched one thus engaged pretty close to me.”

The purpose of this passage is                     .

Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)

As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.

As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.

Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.

Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.

What is the main idea of the passage?

Adapted from The Effects of Cross & Self-Fertilisation in the Vegetable Kingdom by Charles Darwin (1876)

As it is impossible to exclude such minute pollen-carrying insects as Thrips, flowers which it was intended to fertilise with their own pollen may sometimes have been afterwards crossed with pollen brought by these insects from another flower on the same plant; but as we shall hereafter see, a cross of this kind does not produce any effect, or at most only a slight one. When two or more plants were placed near one another under the same net, as was often done, there is some real though not great danger of the flowers which were believed to be self-fertilised being afterwards crossed with pollen brought by Thrips from a distinct plant. I have said that the danger is not great because I have often found that plants which are self-sterile, unless aided by insects, remained sterile when several plants of the same species were placed under the same net. If, however, the flowers which had been presumably self-fertilised by me were in any case afterwards crossed by Thrips with pollen brought from a distinct plant, crossed seedlings would have been included amongst the self-fertilised; but it should be especially observed that this occurrence would tend to diminish and not to increase any superiority in average height, fertility, etc., of the crossed over the self-fertilised plants.

As the flowers which were crossed were never castrated, it is probable or even almost certain that I sometimes failed to cross-fertilise them effectually, and that they were afterwards spontaneously self-fertilised. This would have been most likely to occur with dichogamous species, for without much care it is not easy to perceive whether their stigmas are ready to be fertilised when the anthers open. But in all cases, as the flowers were protected from wind, rain, and the access of insects, any pollen placed by me on the stigmatic surface whilst it was immature, would generally have remained there until the stigma was mature; and the flowers would then have been crossed as was intended. Nevertheless, it is highly probable that self-fertilised seedlings have sometimes by this means got included amongst the crossed seedlings. The effect would be, as in the former case, not to exaggerate but to diminish any average superiority of the crossed over the self-fertilised plants.

Errors arising from the two causes just named, and from others,—such as some of the seeds not having been thoroughly ripened, though care was taken to avoid this error—the sickness or unperceived injury of any of the plants,—will have been to a large extent eliminated, in those cases in which many crossed and self-fertilised plants were measured and an average struck. Some of these causes of error will also have been eliminated by the seeds having been allowed to germinate on bare damp sand, and being planted in pairs; for it is not likely that ill-matured and well-matured, or diseased and healthy seeds, would germinate at exactly the same time. The same result will have been gained in the several cases in which only a few of the tallest, finest, and healthiest plants on each side of the pots were measured.

Kolreuter and Gartner have proved that with some plants several, even as many as from fifty to sixty, pollen-grains are necessary for the fertilisation of all the ovules in the ovarium. Naudin also found in the case of Mirabilis that if only one or two of its very large pollen-grains were placed on the stigma, the plants raised from such seeds were dwarfed. I was therefore careful to give an amply sufficient supply of pollen, and generally covered the stigma with it; but I did not take any special pains to place exactly the same amount on the stigmas of the self-fertilised and crossed flowers. After having acted in this manner during two seasons, I remembered that Gartner thought, though without any direct evidence, that an excess of pollen was perhaps injurious. It was therefore necessary to ascertain whether the fertility of the flowers was affected by applying a rather small and an extremely large quantity of pollen to the stigma. Accordingly a very small mass of pollen-grains was placed on one side of the large stigma in sixty-four flowers of Ipomoea purpurea, and a great mass of pollen over the whole surface of the stigma in sixty-four other flowers. In order to vary the experiment, half the flowers of both lots were on plants produced from self-fertilised seeds, and the other half on plants from crossed seeds. The sixty-four flowers with an excess of pollen yielded sixty-one capsules; and excluding four capsules, each of which contained only a single poor seed, the remainder contained on an average 5.07 seeds per capsule. The sixty-four flowers with only a little pollen placed on one side of the stigma yielded sixty-three capsules, and excluding one from the same cause as before, the remainder contained on an average 5.129 seeds. So that the flowers fertilised with little pollen yielded rather more capsules and seeds than did those fertilised with an excess; but the difference is too slight to be of any significance. On the other hand, the seeds produced by the flowers with an excess of pollen were a little heavier of the two; for 170 of them weighed 79.67 grains, whilst 170 seeds from the flowers with very little pollen weighed 79.20 grains. Both lots of seeds having been placed on damp sand presented no difference in their rate of germination. We may therefore conclude that my experiments were not affected by any slight difference in the amount of pollen used; a sufficiency having been employed in all cases.

One of the main points made in the last paragraph is                     .

Adapted from Essays on Early Ornithology and Kindred Subjects by James R. McClymont (1920)

The voyagers named it the Angra de Santa Elena, and it may have been the bay which is now known as St. Helen’s Bay. But it is worthy of note that the G. de Sta. Ellena of the Cantino Chart is laid down in a position which corresponds rather with that of Table Bay than with that of St. Helen’s Bay.

The Portuguese came into contact with the inhabitants of the country adjacent to the anchorage. These people had tawny complexions, and carried wooden spears tipped with horn—assagais of a kind—and bows and arrows. They also used foxes’ tails attached to short wooden handles. We are not informed for what purposes the foxes’ tails were used. Were they used to brush flies away, or were they insignia of authority? The food of the natives was the flesh of whales, seals, and antelopes (gazellas), and the roots of certain plants. Crayfish or ‘Cape lobsters’ abounded near the anchorage.

The author of the roteiro affirms that the birds of the country resembled the birds in Portugal, and that amongst them were cormorants, larks, turtle-doves, and gulls. The gulls are called "guayvotas," but "guayvotas" is probably another instance of the eccentric orthography of the author and equivalent to "gaivotas."

In December the squadron reached the Angra de São Bràs, which was either Mossel Bay or another bay in close proximity to Mossel Bay. Here penguins and seals were in great abundance. The author of the roteiro calls the penguins "sotelycairos," which is more correctly written "sotilicarios" by subsequent writers. The word is probably related to the Spanish "sotil" and the Latin "subtilis," and may contain an allusion to the supposed cunning of the penguins, which disappear by diving when an enemy approaches.

The sotilicarios, says the chronicler, could not fly because there were no quill-feathers in their wings; in size they were as large as drakes, and their cry resembled the braying of an ass. Castanheda, Goes, and Osorio also mention the sotilicario in their accounts of the first voyage of Vasco da Gama, and compare its flipper to the wing of a bat—a not wholly inept comparison, for the under-surface of the wings of penguins is wholly devoid of feathery covering. Manuel de Mesquita Perestrello, who visited the south coast of Africa in 1575, also describes the Cape penguin. From a manuscript of his Roteiro in the Oporto Library, one learns that the flippers of the sotilicario were covered with minute feathers, as indeed they are on the upper surface and that they dived after fish, upon which they fed, and on which they fed their young, which were hatched in nests constructed of fishbones. There is nothing to cavil at in these statements, unless it be that which asserts that the nests were constructed of fishbones, for this is not in accordance with the observations of contemporary naturalists, who tell us that the nests of the Cape Penguin (Spheniscus demersus) are constructed of stones, shells, and debris. It is, therefore, probable that the fishbones which Perestrello saw were the remains of repasts of seals.

Seals, says the roteiro, were in great number at the Angra de São Bràs. On one occasion the number was counted and was found to be three thousand. Some were as large as bears and their roaring was as the roaring of lions. Others, which were very small, bleated like kids. These differences in size and in voice may be explained by differences in the age and in the sex of the seals, for seals of different species do not usually resort to the same locality. The seal which formerly frequented the south coast of Africa—for it is, I believe, no longer a denizen of that region—was that which is known to naturalists as Arctocephalus delalandii, and, as adult males sometimes attain eight and a half feet in length, it may well be described as of the size of a bear. Cubs from six to eight months of age measure about two feet and a half in length. The Portuguese caught anchovies in the bay, which they salted to serve as provisions on the voyage. They anchored a second time in the Angra de São Bràs in March, 1499, on their homeward voyage.

Yet one more allusion to the penguins and seals of the Angra de São Bràs is of sufficient historical interest to be mentioned. The first Dutch expedition to Bantam weighed anchor on the 2nd of April, 1595, and on the 4th of August of the same year the vessels anchored in a harbor called "Ague Sambras," in eight or nine fathoms of water, on a sandy bottom. So many of the sailors were sick with scurvy—"thirty or thirty-three," said the narrator, "in one ship"—that it was necessary to find fresh fruit for them. "In this bay," runs the English translation of the narrative, "lieth a small Island wherein are many birds called Pyncuins and sea Wolves that are taken with men’s hands." In the original Dutch narrative by Willem Lodewyckszoon, published in Amsterdam in 1597, the name of the birds appears as "Pinguijns."

What is the main idea of the first paragraph?

Adapted from Essays on Early Ornithology and Kindred Subjects by James R. McClymont (1920)

The voyagers named it the Angra de Santa Elena, and it may have been the bay which is now known as St. Helen’s Bay. But it is worthy of note that the G. de Sta. Ellena of the Cantino Chart is laid down in a position which corresponds rather with that of Table Bay than with that of St. Helen’s Bay.

The Portuguese came into contact with the inhabitants of the country adjacent to the anchorage. These people had tawny complexions, and carried wooden spears tipped with horn—assagais of a kind—and bows and arrows. They also used foxes’ tails attached to short wooden handles. We are not informed for what purposes the foxes’ tails were used. Were they used to brush flies away, or were they insignia of authority? The food of the natives was the flesh of whales, seals, and antelopes (gazellas), and the roots of certain plants. Crayfish or ‘Cape lobsters’ abounded near the anchorage.

The author of the roteiro affirms that the birds of the country resembled the birds in Portugal, and that amongst them were cormorants, larks, turtle-doves, and gulls. The gulls are called "guayvotas," but "guayvotas" is probably another instance of the eccentric orthography of the author and equivalent to "gaivotas."

In December the squadron reached the Angra de São Bràs, which was either Mossel Bay or another bay in close proximity to Mossel Bay. Here penguins and seals were in great abundance. The author of the roteiro calls the penguins "sotelycairos," which is more correctly written "sotilicarios" by subsequent writers. The word is probably related to the Spanish "sotil" and the Latin "subtilis," and may contain an allusion to the supposed cunning of the penguins, which disappear by diving when an enemy approaches.

The sotilicarios, says the chronicler, could not fly because there were no quill-feathers in their wings; in size they were as large as drakes, and their cry resembled the braying of an ass. Castanheda, Goes, and Osorio also mention the sotilicario in their accounts of the first voyage of Vasco da Gama, and compare its flipper to the wing of a bat—a not wholly inept comparison, for the under-surface of the wings of penguins is wholly devoid of feathery covering. Manuel de Mesquita Perestrello, who visited the south coast of Africa in 1575, also describes the Cape penguin. From a manuscript of his Roteiro in the Oporto Library, one learns that the flippers of the sotilicario were covered with minute feathers, as indeed they are on the upper surface and that they dived after fish, upon which they fed, and on which they fed their young, which were hatched in nests constructed of fishbones. There is nothing to cavil at in these statements, unless it be that which asserts that the nests were constructed of fishbones, for this is not in accordance with the observations of contemporary naturalists, who tell us that the nests of the Cape Penguin (Spheniscus demersus) are constructed of stones, shells, and debris. It is, therefore, probable that the fishbones which Perestrello saw were the remains of repasts of seals.

Seals, says the roteiro, were in great number at the Angra de São Bràs. On one occasion the number was counted and was found to be three thousand. Some were as large as bears and their roaring was as the roaring of lions. Others, which were very small, bleated like kids. These differences in size and in voice may be explained by differences in the age and in the sex of the seals, for seals of different species do not usually resort to the same locality. The seal which formerly frequented the south coast of Africa—for it is, I believe, no longer a denizen of that region—was that which is known to naturalists as Arctocephalus delalandii, and, as adult males sometimes attain eight and a half feet in length, it may well be described as of the size of a bear. Cubs from six to eight months of age measure about two feet and a half in length. The Portuguese caught anchovies in the bay, which they salted to serve as provisions on the voyage. They anchored a second time in the Angra de São Bràs in March, 1499, on their homeward voyage.

Yet one more allusion to the penguins and seals of the Angra de São Bràs is of sufficient historical interest to be mentioned. The first Dutch expedition to Bantam weighed anchor on the 2nd of April, 1595, and on the 4th of August of the same year the vessels anchored in a harbor called "Ague Sambras," in eight or nine fathoms of water, on a sandy bottom. So many of the sailors were sick with scurvy—"thirty or thirty-three," said the narrator, "in one ship"—that it was necessary to find fresh fruit for them. "In this bay," runs the English translation of the narrative, "lieth a small Island wherein are many birds called Pyncuins and sea Wolves that are taken with men’s hands." In the original Dutch narrative by Willem Lodewyckszoon, published in Amsterdam in 1597, the name of the birds appears as "Pinguijns."

One of the main points made in the last paragraph is                     .

Adapted from "Taking a Second Look: An Analysis of Genetic Markers in Species Relatedness" by Joseph Ritchie (2014)

Phylogenetics is the study of genetic composition in various species and is used by evolutionary biologists to investigate similarities in the molecular sequences of proteins in varying organisms. The amino acid sequences that build proteins are used to construct mathmatical matrices that aid in determining evolutionary ties through the investigation of percentage similarities. The study of these matrices helps to expose evolutionary relationships between species that may not have the same overt characteristics.

Species adapt and evolve based on the pressures that exist in their environment. Climate, food source, and habitat availability are only a few factors that act on species adaptation. These stressors can alter the physical characteristics of organisms. This divergence in evolution has made it difficult to determine the interrelatedness of organisms by analyzing their physical characteristics alone.

For instance, looking only at physical characteristics, the ghost bat resembles a pigeon more than a spider monkey; however, phylogenetics has found that the amino acid sequences that construct the beta hemoglobin molecules of bats are twenty percent more similar to those of mammalian primates than those of birds. This helps reject the assumption that common physical characteristics between species are all that is needed to determine relatedness.

The differences produced by divergent evolution observed in the forest-dwelling, arboreal spider monkey and the nocturnal, airborne ghost bat can be reconciled through homology. Homologous characteristics are anatomical traits that are similar in two or more different species. For instance, the bone structure of a spider monkey’s wrist and fingers greatly resembles that of a bat’s wing or even a whale’s fin. These similarities are reinforced by phylogenetic evidence that supports the idea that physically dissimilar species can be evolutionarily related through anatomical and genetic similarities.

The central idea of this passage is best described by which of the following statements?